Labour overturned its largest ever majority at a by-election – a 24 point swing – to win Selby and Ainsty.
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And in Somerton and Frome the Lib Dems managed a 29 point swing to swoop up the constituency of disgraced ex-MP David Warburton.
In a major blow to Sir Keir Starmer, a huge anti-ULEZ vote helped Tory candidate Steve Tuckwell defy the polls and cling victory.
Conservative chairman Greg Hands said: “An amazing result in Uxbridge!
“It shows that when Labour run something (like London) they run it badly and Sir Keir Starmer flip flops on ULEZ, showing he can’t be trusted.”
An opposition spokesperson admitted Sadiq Khan’s controversial policy was a “concern for voters” but stopped short of calling for it to be scrapped.
In the South West an overjoyed Lib Dem Leader Sir Ed Davey declared his party are “firmly back in the West Country.”
He said: “The people of Somerton and Frome have spoken for the rest of the country who are fed up with Rishi Sunak’s out-of-touch Conservative government.”
Selby will now welcome the first ever MP to parliament born after Tony Blair won in 1997.
25-year-old Keir Mather will replace Boris ultra-loyalist Nigel Adams.
Discussing the victory, Veterans Minister Johnny Mercer last night said: “We don’t want parliament to become like the Inbetweeners.”
Shadow Cabinet Minister Baroness Jenny Chapman hit back: “He’s a young person but he’s not somebody who’s full of themselves.
“He’s intelligent and he’s done a great job.”